1979
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Master in Ecology (Diploma), University of Marburg, Germany |
1981
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PhD, University of Marburg, Germany |
1983/84
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Visiting scientist, Smithsonian Tropical Research Institute, Panama |
1985
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Habilitation in Zoology, University of Marburg, Germany |
1985 - 1988
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Assistant professor, Institute of Zoology, University of Regensburg, Germany |
1988 - 2019
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Full professor for ecology at the Institute of Ecology and Evolution, University of Bern, Switzerland |